In their seminal 1979 paper, evolutionary biologists Stephen Jay Gould and Richard Lewontin introduced the architectural concept of 'spandrels' to evolutionary theory. A spandrel—the triangular space formed by the intersection of two arches—is an inevitable byproduct of structural engineering, not a feature designed for a specific purpose. Similarly, Gould and Lewontin argued, many biological traits are not direct products of natural selection adaptive for survival. Rather, they are structural byproducts of other evolutionary changes. While critics initially dismissed this framework as a mere semantic quibble that undermined the primacy of Darwinian adaptation, the concept eventually revolutionized evolutionary biology. By forcing researchers to distinguish between traits that are direct adaptations and those that are merely secondary consequences of other developments, the spandrel framework redirected the field's focus.
The primary rhetorical function of the second sentence ('A spandrel... specific purpose') is to:
- Ademonstrate that biological evolution is constrained by the same physical laws that govern human architecture.
- Bintroduce a historical counterargument to the idea that biological adaptations are always designed for a specific survival purpose.
- define a structural concept in architecture to serve as an illustrative analogy for the biological theory presented in the passage.Answer
- Dexplain how the literal meaning of a term evolved as it transitioned from architectural engineering to evolutionary biology.