Read the following excerpt regarding ecological succession:
"In the classical model of primary ecological succession, the colonization of barren terrain proceeds through a highly predictable sequence of seral stages. The process invariably commences with autotrophic pioneer species, such as lichens and mosses, which physically and chemically weather the sterile substrate to generate rudimentary soil. Following this pedogenic phase, early successional vascular plants—typically fast-growing, shade-intolerant herbaceous species—colonize the habitat. These r-selected opportunists eventually alter the microclimate by increasing soil moisture and nutrient density, thereby facilitating the establishment of intermediate, shade-tolerant shrubs. The succession theoretically culminates in a self-replacing climax community dominated by K-selected canopy trees. However, modern ecologists note that the ecosystem is rarely immune to allogenic (external) disturbances. Events such as wildfires, extreme meteorological shifts, and anthropogenic interference frequently reset the successional clock."
Statement to evaluate: According to the passage, early successional vascular plants are characterized by several specific traits, but they are NOT described as being shade-tolerant.
Answer: Answer