In examining the perturbation of high-alpine lentic ecosystems in the Sierra Nevada, biologists have elucidated a definitive trophic cascade triggered by the anthropogenic introduction of non-native salmonids. The primary mechanism of this disruption stems from the salmonids' selective predation on large cladoceran zooplankton, notably *Daphnia*. The extirpation of these robust grazers effectively abolishes the competitive suppression they typically exert on smaller, less efficient herbivorous zooplankton. Released from this constraint, the smaller zooplankton proliferate but fail to adequately control phytoplankton biomass. The subsequent exponential proliferation of phytoplankton engenders intense algal blooms that profoundly attenuate the penetration of solar radiation through the water column. This induced turbidity fundamentally alters the benthic habitat by inhibiting the photosynthesis of epilithic algae, thereby depleting the foundational nutritional resource upon which indigenous macroinvertebrate assemblages depend.
Based on the passage, arrange the following phenomena in the chronological order in which they occur as part of the causal chain initiated by the introduction of non-native salmonids.
- 1A precipitous reduction in the population of large cladocerans.
- 2A relaxation of competitive constraints on inefficient herbivorous zooplankton.
- 3An uncontrolled expansion of phytoplankton biomass.
- 4The attenuation of solar radiation reaching the benthic zone.
- 5A depletion of the primary nutritional resources for indigenous macroinvertebrates.